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Tom and Jerry have wrecked the house they lived in during a frantic Tom-Jerry-esque chase, so they enter a race/reality show titled the "Fabulous Super Race", which offers the race-winner a luxurious mansion. Tom and Jerry, having built their own customized vehicles from scrap materials in a junk yard, present themselves to Globwobbler Studios, who are hosting the race. Race commentators are Biff Buzzard and Buzz Blister. The two then meet the contestants: Steed Dirkly, Grammy, Gorthan the Destroyer of Light, Mallory "Soccer Mom" Macdoogle, and Dr. Professor. Dr. Professor is eliminated before the race after Biff touches a Do Not Touch button, causing him to be vaporized.
The racers begin the race in Hollywood, California. When they encounter traffic, all the racers, except Soccer Mom, move up their cars in order to cross. Tom chases Jerry until he sees the tunnel where he falls off the road, causing the other racers to hit him. The race was initially going to be from Hollywood to Mexico, but due to public ratings of the event, Globwobbler decides to drive the racers to the Amazonian, where the finish line is. During the leg, Tom switches a sign, causing contestant Mallory "Soccer Mom" Macdoogle to drive her car into a pit of quicksand. She initially tries to escape using a computer rescue help system, but the system takes too long and she gets out of the car using its open sunroof, before venturing into the jungle, not to be seen for the rest of the movie. Tom then attempts to destroy the bridge, in an attempt to eliminate Jerry. However, Jerry gets to the end of the bridge and Tom's car is set on reverse and falls into a piranha-infested river. However, he gets back up with a contraption in his car. Then J.W. changes the leg to get more ratings. The hosts, Biff and Buzz, then announce the next leg of the race is set in Antarctica, and they will have to modify their cars for Ocean travel.
On the way there, a lightning bolt hits contestant Steed Dirkly's car, causing it to sink. He spots a mermaid on an island, but it turns out she is a monster and feeds Steed to her offspring. The first contestant to reach destination grounds is Gorthan, "Destroyer of Light". However, in a dare by the hosts to test the "your tongue sticks to a cold metal pole" trick (the "triple dog dare" trick from A Christmas Story), his tongue gets stuck and his portion of an ice breaks, eliminating him and leaving him floating away off Antarctica, but not before cursing Biff and Buzz for the trick. Before the next leg of the race, Tom, Jerry and Grammy arrive on the scene and survived the storm. They start to modify their cars, and in a way of sabotoge Tom throws contestant Grammy's dog, Squirty into a whale's mouth and she follows inside and is eaten, supposedly eliminating and killing her. During the leg, Tom and Jerry, the only remaining racers for now, race each other across Antarctica. Meanwhile, in the production office, the producer is notified that the ratings of the race are up and they need something to keep them up. The hosts are then notified that the racers will have to modify their cars to race underwater to Australia. Tom later crashes into a block of concrete and an anchor falls on top his car, supposedly eliminating him.
The movie cuts to Biff and Buzz in Australia with a boxing kangaroo on a leash, though it attacks, fights and beats up Biff, and soon after, Jerry arrives and gestures he has not seen "the pussycat" (Tom). Buzz believes he was accidentally killed in the race. Jerry then prepares to continue racing to Borneo, where J.W. has assigned the finish line to be. Also, Grammy is brought back when the whale spits her out into point. Tom is then brought back up with the assistant of the producer flying to Australia to give him CPR. Tom then repairs his car and cuts the continent in half using a laser, in an attempt to get into first place. However, angry because Tom has destroyed half, the boxing kangaroo comes and attacks Tom. Jerry and Grammy are able to survive.
The next leg of the race involves them modifying their cars for air travel to Borneo, which all three do with balloons, causing them to fly through the air slowly. However, Tom pops Grammy's balloon with a harpoon, causing their car to fall. Grammy and Squirty then fight over a parachute, which turns out to be a lunch bag, causing them to fall to their death. However, Tom pops most of his balloons and Grammy and Squirty's spirits then come up and break the last one, causing him and his car to fall. However, Tom lands on a hammock while Spike is in it, showing where the bulldog was after he ran off. Tom and gets back into his car which lands on Spike who was about to attack him with Jerry following him down to the island. They then race to the finishing point of the leg. In the production office, the producers announce that the final leg of the race back to Hollywood will involve them travelling around The World back to the finishing point in only five minutes, due to the race taking too long. Irving says it may be due to the racers going through the prettiest cities, but J.W. says people don't care about cities, they want action and "plenty of explosions". They decide they will cut the time to five minutes to finish the race. Tom and Jerry are shocked at this, meaning they have only five minutes to go to the opposite side of the world. But they decide to do it anyway for the mansion and the time restrictions. They hop in and race off to make history. The race back involves them completing a path to Asia, then Europe, The Atlantic, and then to Hollywood, from Borneo. At the finish line, Biff and Buzz are waiting for them. Buzz soon witnesses them coming and takes the binoculars from Biff while Biff is still wearing them and nearly chokes Biff to death. Tom and Jerry's jet planes break down at the finish line and move across the finish line at the same time making the race it a tie.
Although they both win because of the tie, the producer says according to their contract, since they tied, they have to do the race all over again. Enraged, Tom and Jerry leap onto J.W. and fight and beat him up for fact they could've been killed during the race. Irving watches in shick as the animals use anvils, mousetraps and sledgehammers against J.W. At the end, J.W. tries to climb out of the skirmish by climbing on the Finish line strip, but Tom and Jerry grab him and bring him down. They take the key to the mansion from him like The Battle and the Furry. The producer, angry and disoriented, decides to change Hollywood's focus to family programming. The main boss appears and kills him by cremating him for having a family friendly attitude. The main boss, subsequently makes the producer's assistant the head of Globwobbler studios. At last, all is well with Tom and Jerry now sharing the mansion peacefully until the owner of their previous house shows up and orders Tom to get rid of Jerry. The movie ends with Tom chasing Jerry around their new home, presumably going to be destroyed again.
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